KEY TO THE FAMILIES OF HARPACTICOIDA

(amended from Lang, 1948)

 

1.

Commensal on whales.

Balaenophilidae a)

 

Not commensal on whales.

2

2.

Body laterally compressed, as Fig. 4.

Tegastidae

 

Body dorso-ventrally compressed, with the expanded epimera of the anterior abdominal segments enfolding the posterior abdomen and caudal furca, giving the body the shape of a wood-louse. (Figs. 5-6).

3

 

Body of a different shape from either of above conditions.

4

3.

Body-shape as Fig. 5; without clearly seen chitin “skeleton”; P.1 as Fig. 22.

Porcellidiidae a)

 

Body-shape as Fig. 6; a chitinous “skeleton” clearly visible; P .1 as Fig. 23.

Peltidiidae

4.

Exp. A.2 of at least 6 segments.

5

 

Exp. A.2 of not more than 4 segments.

6

5.

Distal segment of Enp. P.2 much longer than the entire Exp.

Longipediidae a)

 

Distal segment of Enp. P.2 shorter than the Exp.

Canuellidae

6.

Segment bearing the P.1 not fused to the cph.; Exp. A.2 of 1 segment with several setae; Mxp. lamelliform with several setae. Mosses and freshwaters.

Phyllognathopodidae a)

 

Segment bearing the P.1 not fused to the cph.; Exp. A.2 of 2 segments; Mxp. not lamelliform. Freshwaters.

Chappuisiidae a)

 

Segment bearing the P.1 only partially fused with the cph.; Exp. A.2 absent or represented by 1-2 setae only; Mxp. degenerate; Fresh and salt waters.

Darcythompsoniidae

 

Segment bearing the P.1 completely fused with the cph.

7

7.

Mx2 as Figs. 13-15; body fusiform or vermiform; Benp. P.5 with only 2 setae (Benp. and Exp. of P.5 may be fused together but the parts are always recognizable) (Fig. 36).

Ectinosomatidae

 

These characters not combined.

8

8.

P.1 as Fig. 24.

Harpacticidae

 

P.1 as Fig. 25.

Clytemnestridae a)

 

P.1 as Fig. 26 (Enp. may be 2 or 3 segments).

Tisbidae

 

P.1 as Fig. 27.

Tachidiidae

 

P.1 as Fig. 28 (Exp. can be 1 to 3 segments; 2nd Enp. segment may bear 1 claw or two long setae); Exp. A.2 present at least as a rudiment.

Laophontidae

 

P.1 as Fig. 29; Body pyriform; mouth-organs degenerate.

Metidae

 

Exp. P.1 absent or of 1 small segment; Enp. of 2 segments (Figs. 30-31).

Cylindropsyllidae

 

P.1 not as any of the above, or, if similar to Fig. 28 then Exp. A.2 absent.

9

9.

Exp. P.1 of 3 segments, middle segment without a spine on the outer border.

10

 

Exp. P .1 of 3 segments, middle segment with a spine on outer border or Exp. P.1 of less than 3 segments.

12

10.

Enp. P.2-P.4 of 1 segment; Exp. P.3 (M) modified; 2nd segment Exp. P.1 without an inner seta.

Parastenocarididae a)

(freshwater)

 

Enp. P.2-P.4 of 1:1:2 segments respectively; Exp. P.3 (M) not modified; 2nd segment Exp. P.1 without an inner seta.

Cylindropsyllidae

 

Enp. P.2-P.4 of 2:1:2 segments respectively; Exp. P.3 (M) not modified; 2nd segment Exp. P.1 with an inner seta.

Cylindropsyllidae

 

Enp. P.2-P.4 each of 2 segments; Exp. P.3 (M) not modified; 2nd segment Exp. P.1 with or without an inner seta.

11

11.

2nd segment Exp. P.1 with an inner seta.

Ameiridae

 

2nd segment Exp. P.1 without an inner seta.

Cylindropsyllidae

12.

1st segment Exp. P.1 with an inner seta; Mxp. not prehensile.

13

 

These characters not combined.

14

13.

Exp. A.2 of 1 segment, with at most 2 setae.

Aegisthidae a)

 

Exp. A.2 of 3-4 segments with several setae.

Cerviniidae

14.

Enp. P.4 at most 2 segments.

15

 

Enp. P.4 of 3 segments.

31

15.

Exp. A.2 of 4 segments; Mxp. not prehensile; Enp. P.1-P.4 with 3:2:2:2 segments respectively.

Neobradyidae a)

 

Exp. A.2 of 3 segments.

16

 

Exp. A.2 at most 2 segments.

17

16.

Mxp. not prehensile; Enp. P.1-P.4 with 2:3:2:2 segments respectively.

Tisbidae

 

Mxp. prehensile; Enp. P.1-P.4 each of 2 segments.

Diosaccidae

17.

Mxp. not prehensile; A.1 (F) of 5-6 segments with plumose setae, all segments serrated dorsally.

Tisbidae

 

Mxp. not prehensile; remainder of above characters not combined.

18

 

Mxp. prehensile.

20

18.

P.4 as Fig. 33 (Enp. may be 1 or 2 segments); P.5 a single small plate; A.1 (F) of not more than 7 segments.

Cylindropsyllidae

 

These characters not combined.

19

19.

Distal segment Exp. P.2-P.4 with 3 spines on the outer border.

Louriniidae a)

 

Distal segment Exp. P.2-P.4 with at most 2 setae and/or spines on the outer border.

Darcythompsoniidae

20.

Mxp. as Fig. 20.

Cylindropsyllidae

 

Mxp. not as above.

21

21.

Exp. P.4 as Fig. 33 (Enp. may be 1 or 2 segments); P.5 a single plate.

Cylindropsyllidae

 

These characters not combined.

22

22.

Mxp. as Fig. 19.

Paramesochridae

 

Mxp. not as above.

23

23.

Labrum large and strongly prominent; Rostrum articulated with cph.; Cu.f. always distinctive in shape.

Tetragonicipitidae

 

These characters not combined.

24

24.

Exp. A.2 absent; A.1 (F) at most 5 segments.

Ancorabolidae

 

These characters not combined.

25

25.

All body segments sharply differentiated; Enp. P.1 not prehensile, at most of 2 segments.

Cletodidae

 

These characters not combined.

26

26.

A.1 (F) of 4-5 segments; Enp. P.1 not prehensile.

Cletodidae

 

These characters not combined.

27

27.

P.1 as Fig. 32.

Ameiridae

 

P.1 not as above; basis of P.2-P.4 transversely prolongated.

Ameiridae

 

P.1-P.4 not as either of above.

28

28.

A.2 with basis (Fig. 8).

Ameiridae

 

A.2 with allobasis (Fig. 9).

29

29.

Rostrum large; P.5 as Figs. 37-38.

Diosaccidae

 

Rostrum large; P.5 not as above; A.1 (F) of 4 segments; Exp. A.2 of 1 segment.

Canthocamptidae

 

Not as either above.

30

30.

Enp. P.1 prehensile; Enp. P.2-P.3 of 3 segments.

Diosaccidae

 

These characters not combined.

Canthocamptidae

31.

Cuticular lenses present.

Miraciidae a)

 

No cuticular lenses.

32

32.

Body shape as Fig. 7.

Miraciidae a)

 

Body shape not as above.

33

33.

A.2 with basis (Fig. 8); Exp. A.2 of 1 segment; Mx2 with at most 3 endites.

34

 

These characters not combined.

38

34.

P.5 with Benp. and Exp. distinct.

35

 

P.5 with Benp. and Exp. fused to form a single plate.

37

35.

Enp. P.1 of 2 segments, not prehensile.

Cletodidae

 

Enp. P.1 of 3 segments.

36

36.

1st segment Exp. P.1 with an inner seta.

Cletodidae

 

1st segment Exp. P.1 without an inner seta.

Ameiridae

37.

Distal segment Exp. P.2-P.4 each with at least 5 setae and/or spines.

Ameiridae

 

Distal segment Exp. P.2-P.4 each with only 4 setae and/or spines.

Latiremidae a)

 

Distal segment Exp. P.2-P.4 with 4:5:5 setae and/or spines respectively.

Cletodidae

38.

Mxp. well developed, not prehensile.

Tisbidae

 

Mxp. not as above.

39

39.

Enp. P.1 not prehensile.

40

 

Enp. P.1 prehensile.

43

40.